David Clark





Educational Background

  • Ph.D., University of Cincinnati (1992)
  • M.S., Central Michigan University (1987)
  • B.S., Central Michigan University (1982)



My career at Alma began in


I'm an expert in

animal communication and evolution

My expertise:

Dr. Clark’s research focuses upon animal communication and the evolution of visual displays. Actively engaging undergraduate students in his research, he studies a variety of animals including spiders, lizards, fish and turtles. In 2012, Clark was the visiting scientist on site with British broadcaster and naturalist Sir David Attenborough on a 3D documentary, The Galapagos 3D. The documentary included a segment featuring Clark’s research using robotic lizards to test questions about lizard signaling and communication.

Recent publications:

Gillingham, J.C. and Clark, D.L. (In Press) Normal Behavior. Ed. G. Burghardt and C. Warwick, in Health and Welfare of Captive Reptiles, Springer Press

Clark, D.L., Macedonia, J.M., Rowe, J.W., Austin, M.R., Centurione, I.M., Valle, C.A. (2019 Review). Galápagos Lava Lizards (Microlophus bivittatus) Respond Dynamically to Displays from Interactive Conspecific Robots. Behavioral Ecology and Sociobiology 73:136-146

Uetz, G.W., Clark, D.L., Kane, H., Stoffer, B. 2019. Listening in: the importance of vibratory courtship signals for male eavesdropping in the wolf spider, Schizocosa ocreata. Behavioral Ecology and Sociobiology 72:133-141.

Macedonia, J.M., Clark, D.L., Fonley, M.R., Centurione, I., Rowe, J.W., Valle, C.A. (In Press). Analysis of Bobbing Displays in Four Species of Galápagos Lava Lizards Using Conventional and Novel Quantitative Methods, Herpetologica.

Rubi, T.L., Clark, D.L., Keller, J.S., Uetz, G.W. 2019. Courtship behavior and coloration influence conspicuousness of wolf spiders (Schizocosa ocreata (Hentz)) to avian predators. Behavior Processes 162:215-220

Lietzenmayer, L.B., Clark, D.L., Taylor, L.A. (In press). The role of male coloration and ornamentation in potential alternative mating strategies of the dimorphic jumping spider, Maevia inclemens. Behavioral Ecology.